Human Genetics and Genomics, PhD

School of medicine, ph.d. program.

The Johns Hopkins Human Genetics Training Program provides a training in all aspects of human genetics and genomics relevant to human biology, health and disease. 

Advances in human genetics and genomics continue at an astounding rate and increasingly they are being integrated into medical practice. The Human Genetics Program aims to educate highly motivated and capable students with the knowledge and experimental tools that will enable them to answer important questions at the interface between genetics and medicine. Ultimately, our trainees will be the leaders in delivering the promise of genetics to human health.

The overall objective of the Human Genetics program is to provide our students with a strong foundation in basic science by exposure to a rigorous graduate education in genetics, genomics, molecular biology, cell biology, biochemistry and biostatistics as well as a core of medically-related courses selected to provide knowledge of human biology in health and disease. 

This program is also offered as training for medical students in the combined M.D./Ph.D. program.  Students apply to the combined program at the time of application to the M.D. program. (See section entitled Medical Scientist Training Program).

Research Facilities

Research laboratories are well equipped to carry out sophisticated research in all areas of genetics. The proximity to renown clinical facilities of the Johns Hopkins Hospital, including the Department of Genetic Medicine, and Oncology Center provides faculty and students with access to a wealth of material for study. Computer and library facilities are excellent. Laboratories involved in the Human Genetics Program span Johns Hopkins University; consequently supporting facilities are extensive.

Financial Aid

The program is supported by a training grant from the National Institute of General Medical Sciences. These fellowships, which are restricted to United States citizens and permanent United States residents, cover tuition, health care insurance and a stipend during year one.  Once a student has joined a thesis lab, all financial responsibilities belong to the mentor.   Students are encouraged, however, to apply for fellowships from outside sources (e.g., the National Science Foundation, Fulbright Scholars Program, Howard Hughes Medical Institute) before entering the program.

Applicants for admission should show a strong academic foundation with coursework in biology, chemistry and quantitative analysis.   Applicants are encouraged to have exposure to lab research or to data science.  A bachelor's degree from a qualified college or university will be required for matriculation.  GREs are no longer required.

The Human Genetics site has up-to-date information on “ How to Apply .” For questions not addressed on these pages, please access the contact imformation listed on the program page: Human Genetics and Genomics Training Program | Johns Hopkins Department of Genetic Medicine (hopkinsmedicine.org) .

Program Requirements

The program includes the following required core courses: Advanced Topics in Human Genetics, Evolving Concept of the Gene, Molecular Biology and Genomics, Cell Structure and Dynamics, Computational Bootcamp,  Pathways and Regulation, Genomic Technologies, Rigor and Reproducibility in Research, and Systems, Genes and Mechanisms of Disease. Numerous elective courses are available and are listed under sponsoring departments.

Our trainees must take a minimum of four electives, one of which must provide computational/statistical training.

The HG program requires the “OPTIONS” Career Curriculum offered by the Professional Development and Career Office.  OPTIONS is designed to provide trainees with the skills for career building and the opportunity for career exploration as well as professional development training

Human Genetics trainees also take a two-week course in July at the Jackson Labs in Bar Harbor, Maine entitled "Human and Mammalian Genetics and Genomics: The McKusick Short Course" which covers the waterfront from basic principles to the latest developments in mammalian genetics. The faculty numbers about 50 and consists roughly in thirds of JAX faculty, Hopkins faculty and “guest” faculty comprising outstanding mammalian geneticists from other US universities and around the world.

The courses offered by the faculty of the program are listed below. All courses are open to graduate students from any university program as well as selected undergraduates with permission of the course director.

Trainees must complete three research rotations before deciding on their thesis lab.  They must also participate in the Responsible Conduct of Research sessions offered by the Biomedical Program; starting at year 3, students must attend at least two Research Integrity Colloquium lectures per year. 

Our trainees participate in weekly journal clubs, department seminars, monthly Science & Pizza presentations as well as workshops given twice a year on diversity, identity and culture.

At the end of the second year, trainees take their Doctoral Board Oral Examination.  Annual thesis committee meetings must be held following successful completion of this exam.

Average time for completion is 5.3 years.

Graduates from the Human Genetics program pursue careers in academia, medicine, industry, teaching, government, law, as well the private sector.  Our trainees are encouraged to explore the full spectrum of professional venues in which their training my provide a strong foundation. Driven by curiosity and a desire for excellence, our trainees stand out as leaders in the chosen arenas of professional life. They are supported in the development of their career plans by a program faculty and administration who are dedicated to their success, and by a myriad of support networks across the Johns Hopkins University, many of which are provided by the Professional Development Career Office of the School of Medicine.

Ph.D. Program

The training for a Ph.D. in Biology is focused on helping students achieve their goals of being a successful research scientist and teacher, at the highest level. Students work closely with an established advisor and meet regularly with a committee of faculty members to facilitate their progress. The Biology Ph.D. program is part of the larger Biosciences community at Stanford, which includes doctorate programs in the basic science departments at Stanford Medical School. 

There are two tracks within the Biology Ph.D. program:

  • Cell, Molecular and Organismal Biology
  • Ecology and Evolution

(Previously a part of the Department of Biology Hopkins Marine Station is now a part of the Oceans Department within  Stanford Doerr School of Sustainability )

All  tracks are focused on excellence in research and teaching in their respective areas; where there are differences between the tracks, they are indicated in the links below. 

Requirements & Forms

Dissertation defense, cellular and molecular biology training program, stanford biology ph.d. preview program, career development resources.

Department of Biological Sciences

College of natural and mathematical sciences, phd in molecular and cell biology.

Female student holds pipette while sitting at lab bench.

The intent of the MOCB program is to provide an intensive, innovative but broadly-based training in molecular and cell biology. To assure that students have a proper grounding the program prescribes a set of advanced graduate classes in biochemistry, genetics and molecular biology (prokaryotic and eukaryotic). In addition, students get specific training in some aspect of molecular and cell biology by participating in at least one three-credit graduate seminar and by continued participation in research seminars (or journal clubs) in their area of concentration.

Degree Requirements

The degree requirements include a total of 8 graduate level courses and completion of 12 credits of research:

  • Two-semester course in biochemistry
  • One course based on the principles of genetic analysis
  • One course in prokaryotic molecular biology
  • One course in eukaryotic molecular biology
  • One or two 700-level graduate seminars (included in the 18 credits of 600/700 level courses)
  • Elective courses to bring the total of courses taken to 8
  • 12 credits of research. At least one research credit each semester
  • Beginning with the second year, students are required to participate in a research seminar (journal club) every semester and to take one credit of research seminar (journal club) each year

In addition, students who complete the degree will have completed

  • Three (3) research rotations on a short research project lasting 10-12 weeks under the supervision of a faculty member in the program
  • Two semesters as a teaching assistant
  • Most important, the student will complete original research under the direction of their faculty mentor and overseen by a committee knowledgeable in the student’s area of specialization
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EMBL International PhD Programme

Unique in the world and waiting for you!

The EMBL International PhD Programme (EIPP), originally established in 1983, provides PhD students with an excellent starting platform for a successful career in science by  fostering early independence and interdisciplinary research .

EMBL is dedicated to  promoting excellence  in the molecular life sciences throughout Europe. To achieve this goal, we inspire and train talented young scientists to become skilled and creative future leaders in academia, industry and other sectors.

The EIPP provides comprehensive interdisciplinary training, maintaining a careful balance between theory and practice, close mentoring and creative freedom, collaborative teamwork and independence. These key characteristics make the EIPP a  role model , which has inspired similar programmes at research institutions throughout Europe and the world.

The  enriching  encounter of different nationalities, the  friendly and collaborative atmosphere , and the  passion for science  is what unites EMBL’s diverse staff and provides the ideal setting to forge long-lasting connections and make studying at EMBL a unique formative experience.

fully funded phd molecular biology

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fully funded phd molecular biology

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fully funded phd molecular biology

Nationalities represented among PhD students

fully funded phd molecular biology

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fully funded phd molecular biology

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fully funded phd molecular biology

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Application

Two recruitment rounds are organised each year across all EMBL sites. The application deadlines and all relevant information are published well in advance.

Registration for the 2024 PhD Summer Recruitment is now closed.

The next call for applications will open in September 2024.

Programme Overview

Explore the structure and content of the EMBL International PhD Programme.

Research at EMBL

Find out more about the cutting-edge Research Topics investigated across our different Research Units .

The complexity of current biological research questions increasingly calls for interdisciplinary approaches . To prepare our students for this challenge, the EIPP puts great emphasis on bringing together different fields of expertise: we actively encourage candidates with backgrounds in chemistry, physics, mathematics, molecular medicine, computer science or engineering to apply and work closely with biologists to answer fundamental questions in life sciences.

PHD Symposium

Since its first edition in 2000, the annual EMBL International PhD Symposium in Heidelberg developed into a well-recognized and highly appreciated scientific event.

It is always EMBL’s first year PhD students who take on the responsibility for the conceptual development and organization of the meeting. The symposia provide a great platform for networking and scientific exchange, featuring lectures by top-level scientists as well as plenty of opportunities for young researchers to present their work. 

See the full list of past symposia

fully funded phd molecular biology

Upcoming symposium

Power of Many: Collective Behaviour Across Scales

20 – 22 November 2023 EMBL Heidelberg

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Project and Programme Manager

Postdoctoral Programme

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EMBL Graduate Office

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Tel: +49 6221 387-8612/8896 Fax: +49 6221 387-8400 Email:   [email protected]

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Program components, m.d./ph.d. programs.

  • VAI-MSU Graduate School Program

One Degree. Endless Possibilities.

Van Andel Institute Graduate School’s Ph.D. in molecular and cellular biology is a research-intensive, interdisciplinary program that prepares students for successful careers as independent investigators. Students receive in-depth training in the latest techniques across fields and disease areas, including epigenetics, genetics, cancer, neuroscience, metabolism, immunology, structural biology, bone biology and more. The Graduate School uses an innovative, problem-based learning approach that trains students to conduct high-caliber, rigorous science and to translate basic biological findings into clinical applications, blending discovery with invention and insight with application. Students also undergo extensive professional development in areas such as leadership, ethics, responsible and effective conduct of research, public speaking, and grant and technical writing. By the end of their time at the Graduate School, students are prepared to tackle science’s most pressing questions.

Our Ph.D. program comprises a hands-on, scientifically rigorous and well-rounded problem-based approach.

See a year-by-year view of Graduate School coursework.

Course Catalog

Our courses are designed to transform the graduate students of today into the biomedical research leaders of tomorrow.

We partner with Michigan State University College of Human Medicine and Western Michigan University Homer Stryker M.D. School of Medicine to offer dual M.D./Ph.D. programs.

Doctoral Components

Designed to prepare students to be successful independent investigators, Van Andel Institute Graduate School’s Ph.D. program comprises a hands-on, scientifically rigorous and well-rounded problem-based approach.

Students complete modules in biochemistry, cell and molecular biology, genetics and epigenetics, bioinformatics, and pathophysiology before selecting the laboratory in which they will complete their dissertation research. This approach allows students to acquire the requisite knowledge to address complex research questions along with the skills to locate and evaluate the concepts, models and evidence that already exist in scientific literature. For specific course descriptions, please view our Academic Catalog .

Ph.D. degree course requirements

The Graduate School’s curriculum is designed to train students to think like scientists through a problem-based approach. Most courses are completed in the first year of study, followed by a heavily research-focused approach during years three through five. For a sample year-by-year schedule, please visit the Curriculum guide .

Research Rotations

New students complete three, four-week rotations in Van Andel Institute laboratories prior to selecting a thesis adviser. These rotations give students in-depth insight into the different types of research underway at the Institute and help identify students’ particular area of interest.

For more information on the Institute’s labs, please visit our Faculty Directory .

Comprehensive Examination

The comprehensive examination is completed in the second year of the Graduate School’s program. Students write a proposal outlining the background, preliminary results and experimental plan for their thesis research. Students also draft a second proposal to explore an additional, non-thesis research question. During an oral examination, faculty members test the student’s knowledge of concepts and methods in their field of study as well as the student’s critical analysis skills.

Thesis Research

The completion of an original and creative research project resulting in the compilation of a doctoral dissertation is the capstone to the Graduate School’s doctoral degree program. The dissertation demonstrates that the student has the knowledge, skill and scientific maturity to design and conduct independent research of a quality consistent with reports published in peer-reviewed scientific journals. A Graduate School faculty member serves as the principal thesis adviser for this research. Other faculty serve on the student’s thesis advisory committee, which meets at least twice annually to review and evaluate progress and plans for dissertation work.

Seminars, Journal Clubs and Research Reports

Graduate School students meet weekly for a seminar that alternates between research in progress reports and presentations on new papers in scientific literature. Students are expected to attend each week and to present at this seminar at least once each year.

Van Andel Institute also convenes a separate weekly seminar on research underway in its labs. These internal presentations are supplemented by the VAI Seminar Series, which bring outstanding scientists from around the world to the Institute to give scientific talks. Students are encouraged to attend these seminars and to meet with the visiting scientists to discuss their research and their scientific career path.

Professional Development

The Graduate School’s curriculum is bolstered by workshops and training sessions to facilitate the transition from graduate student to professional scientist. These events include guidance and training in writing and reviewing grants and manuscripts, oral presentation skills, financial and personnel management, conflict resolution, and leadership and team-building skills. Additionally, each Graduate School student is allowed up to $2,500 annually to attend national or international meetings relevant to their field of study.

Teaching Experience

Given that many scientists serve as faculty at institutions of higher education, training in the teaching of science at the college level is available depending on student interest. Teaching experiences are coordinated with local colleges and universities that offer appropriate courses in which the student can serve as a teaching assistant, with instruction and supervision by experienced faculty.

Curriculum Overview

All Van Andel Institute Graduate School students follow a similar curriculum schedule, with most courses being completed during the student’s first and second years. Years three through five are heavily focused on the student’s dissertation work as well as professional development.

Students complete five, four-week modules in biochemistry, cell and molecular biology, genetics and epigenetics, bioinformatics and pathophysiology before selecting the laboratory in which they will complete their dissertation research. Through this approach, students acquire the requisite knowledge to address complex research questions and the skills to locate and evaluate the concepts, models and evidence that already exist in scientific literature. For a full description of the Graduate School’s offerings, please view our Course Catalog .

fully funded phd molecular biology

Sample Year-by-Year Schedule

fall semester (16 weeks)

Strategic Approaches to Biomedical Research (SABR) Modules (3)

Historical Perspectives in Molecular Biology

Laboratory Rotation (1) *Students select a research adviser after completing these rotations.

Experimental Design and Biostatistics (Professional Development)

Experimental Skills 1 (Professional Development)

Scientific Communication 1 (Professional Development)

Winter semester (17 weeeks including exam)

Strategic Approaches to Biomedical Research (SABR) Modules (2, plus exam)

Responsible and Effective Conduct of Research (Professional Development)

Laboratory Rotation (2 credits each) *Students select a research adviser after completing these rotations.

4 total (2 credits each)

Data Analysis and Bioinformatics (Professional Development)

Experimental Skills 2 (Professional Development)

Scientific Communication 2 (Professional Development)

Summer semester (14 weeks) course

Precandidacy research

Thesis Research (Precandidacy and Doctoral Candidate)

Up to 6/semester

1 or 2 each

Special Topics

Doctoral Candidate Thesis Research

Teaching Opportunity (if desired)

Final Thesis Advisory Committee Meetings(s)

Dissertation Preparation

Dissertation Defense

Course Descriptions

Vai 8010-8051, strategic approaches to biomedical research (sabr).

Fall and Winter Semesters, 11 Credits

In a progressive series of four-week modules, students develop research plans to address current hypotheses, questions or problems relevant to human disease. In the course of developing these plans, students learn foundational concepts in biochemistry, cell biology, molecular biology, genetics, bioinformatics, and pathobiology. This “problem-based learning” approach best simulates how professional scientists attack new research problems. Students emerge with a strong foundation in core concepts in the relevant disciplines, an understanding of experimental design principles, and experience in crafting research plans. Across the Fall and Winter semesters of the first year, students undertake a total of five two-credit SABR modules and a one-credit, cumulative final examination.

Fall Semester, 2 Credits

This course examines the historical context of current molecular and cell biology research. Students study classic papers in biomedical research and discuss how the work represented in those papers changed the models or paradigms that prevailed at the time the research was done. Topics include foundations of modern biology, mechanisms of genetic change, analysis of biological macromolecules, gene splicing and rearrangement, disease mechanisms, tumor suppressor genes, and organisms used as important experimental models.

Responsible and Effective Conduct of Research

Winter Semester, 2 Credits

This course addresses effective laboratory management practices including protection of human and animal subjects, scientific integrity, conflicts of interest, collaboration, authorship, peer review, data management, mentoring, communication, societal impacts, human resource management, grants and contracts, and fiscal responsibility. The course provides training and direction on how to recognize, address and prevent ethical dilemmas that arise during the course of conducting scientific research.

Experimental Design and Biostatistics

An increasing emphasis on rigor and reproducibility has highlighted the fundamental roles of experimental design and statistics in modern biological research. This course focuses on basic principles of experimental design and fundamental statistical concepts for modern data-intensive biological research. The material draws upon methods and applications from concurrent subject-specific modules. Topics include probability, random variables, sampling, estimation, hypothesis testing, linear regression, diagnostics for fit, model selection, and ANOVA. Students will develop skills in R with RStudio.

Data Analysis and Bioinformatics

Many research projects in modern molecular and cell biology require the analysis of very large datasets such as those generated in genomics, epigenetics, metabolomics, proteomics, and structural biology. Almost all aspects of modern biology incorporate large-scale data analysis to some extent. The efficient and accurate analysis and interpretation of these datasets are fundamentally important activities in biomedical research. This course delves into the algorithms and tools used in the application of bioinformatics to high-dimension datasets. Students will expand upon the R skills developed in the Biostatistics course and apply the skills to genomic, epigenomic, transcriptomic, and proteomic datasets, as well as downstream and integrative analysis.

VAI 8260 – 8261

Scientific communication 1 and 2.

Fall and Winter Semesters, 2 Credits Each

This course is intended to help students become more effective communicators in their scientific work. The scientific research process relies heavily on effective communication of concepts, plans, results, and conclusions. For that reason, scientists must be skilled in spoken and written communication. The course will provide foundational principles and iterative practice in communication as listeners, speakers, readers, and writers, in multiple formats and with various audiences. The formats for listening and speaking will include formal scientific presentations, chalk talks, lab meetings, talks for lay audiences, and posters. The written formats will include grant proposals, scientific papers, review articles, and lay summaries. Course content and activities will align with concurrent courses and with laboratory rotation experiences.

VAI 8250 – 8251

Experimental skills 1 and 2.

Fall Semester, 2 Credits; Winter Semester, 1 Credit

This course will provide a focused introduction to well-established and cutting edge technologies, instrumentation, and methods important for addressing the scientific problems explored in Strategic Approaches to Biomedical Research (SABR), with an emphasis on technologies available in the Van Andel Institute's Technologies and Services. The goal is for students to develop the knowledge and critical thinking skills needed to effectively incorporate these well-established and cutting-edge technologies, instrumentation, and methods into their own research.

Technical Writing and Grantsmanship I

Fall Semester, 1 Credit

This course is intended to help students become more effective writers in scientific disciplines. The entire research process depends upon the communication of concepts, results and plans. For that reason, scientists must be skilled in communicating through presentation and in writing. The course addresses the characteristics of clarity, organization, and style in technical writing and especially in scientific proposals. A major theme of the course is the process of writing, involving composition, editing, and revising with feedback. Students participate in multiple exercises with opportunities for review and iterative development of a draft proposal.

VAI 9301-9313

Professional development courses.

1–2 Credits Per Course

These courses build student skills in communication, laboratory management, and organization. Courses complement the External and Internal Seminar Reporting. Recent offerings include Grantsmanship, Lab Leadership, and Origins of Cancer Scientific Conference Organization.

VAI 9001- 9024

Special topics courses.

1–2 credits Per Course

These courses provide advanced study on focused topics in basic biomedical research, and are typically taken in the second, third, and fourth years. Each course engages students in the study and discussion of the current scientific literature and concepts of the topic selected. Specific content varies with each semester. Special topics courses in various fields are offered on a rotating basis. Additional courses may also be offered depending on student and faculty interest.

Laboratory Rotations

Minimum 3 Rotations, 6 Credits

Laboratory rotations in the first year provide early research experiences that are important in the development of students. These laboratory rotations assist students in their choice of a thesis adviser, laboratory, and dissertation project. Students will complete at least three rotations. The activities of the rotation should be planned to give the student a rich and deep understanding of the questions being addressed, the approaches and experimental methods employed, the mentoring and leadership style of the laboratory head, and the relationships with other members of the laboratory team. Students should expect to spend as much time in the laboratory as their course work will allow (typically 30 hours per week).

Independent Study

Credits Vary Depending on Effort

Students request Van Andel Institute Gradate School academic credit for a course or workshop taken at another institution (whether in-person or online), or for learning experiences at Van Andel Institute that are not part of existing courses. Requests are evaluated on a case by case basis.

M.D./Ph.D. Dual-Degree Programs

Van Andel Institute Graduate School partners with Michigan State University College of Human Medicine and Western Michigan University Homer Stryker M.D. School of Medicine to offer training in both clinical medicine and biomedical research, resulting in both M.D. and Ph.D. degrees. These programs typically span eight years, with four years of Ph.D. research training flanked on each side by two years of medical and clinical education.

Van Andel Institute Graduate School and Michigan State University College of Human Medicine

This M.D./Ph.D. program is a joint effort between Van Andel Institute Graduate School and Michigan State University College of Human Medicine. Students admitted to the program will pursue an M.D. degree through Michigan State while simultaneously pursuing a Ph.D. through Van Andel Institute Graduate School.

Students enrolled in this combined program receive their preclinical education and training through the College of Human Medicine and their biomedical research education, including coursework, laboratory research, and dissertation research through Van Andel Institute Graduate School. They are considered students of both institutions for the duration of their participation in the program. Students receive priority placement at the College of Human Medicine’s Grand Rapids campus for the preclinical and clinical training years.

Students spend years one and two completing their College of Human Medicine preclinical coursework and taking the USMLE Step 1 Exam. Beginning in year three, students attend Van Andel Institute Graduate School to complete the Ph.D. portion of the program. Students will be able to apply their Ph.D. research as two College of Human Medicine research electives toward their clerkship requirements during the fourth year, decreasing the duration of medical school by eight weeks. Students are expected to complete the Ph.D. portion of the degree by the end of year five and prior to beginning their clinical clerkships.

Van Andel Institute Graduate School will award the Ph.D. degree when the student completes all of the degree requirements. The student will then return to the College of Human Medicine to complete the final two years of clinical training necessary for the M.D. degree.

During the preclinical and clinical medical school years, students will pay regular Michigan State University M.D. program tuition. They may apply for competitive fellowships and scholarships both internal and external to the College of Human Medicine.

During the graduate school years, students will receive a package of financial assistance administered through Van Andel Institute Graduate School. This support, typically directed through a graduate fellowship, which includes a stipend; health, dental, vision and life insurance; a full waiver of tuition to the Graduate School; and $2,500 a year for travel to scientific conferences. Students will be expected to apply for external predoctoral fellowships, which will be submitted and administered through the Graduate School and relevant Van Andel Institute offices. During the graduate school years, no tuition will be directed or paid to MSU.

External training grants to support physician-scientist training programs are or may become available from federal, state or private sources. Applications for such training grants will be administered through Michigan State College of Human Medicine. Van Andel Institute Graduate School will be a cooperating institution on such training grants. Funds supporting students (stipends, benefits, tuition relief, etc.) during their medical school years will be directed to Michigan State, and funds supporting students during their graduate school years will be directed to Van Andel Institute Graduate School.

Application and Eligibility

An applicant must apply for admission to MSU College of Human Medicine through the American Medical College Application Service (AMCAS) as a Combined Medical Degree/Ph.D. applicant. Applicants do not need to fill out a separate Van Andel Institute Graduate School application; however, to apply to the joint M.D./Ph.D. program, applicants must check the appropriate box on the AMCAS application.

If the applicant is deemed eligible for acceptance by both the College of Human Medicine and Van Andel Institute Graduate School, the applicant will be reviewed by the joint M.D./Ph.D. Selection Committee for a final decision. If accepted by the M.D./Ph.D. Selection Committee, the applicant will be offered a position in the M.D./Ph.D. program.

Acceptance to the College of Human Medicine M.D. Program is contingent upon acceptance by the M.D./Ph.D. Program Selection Committee.

Interested applicants should:

  • Apply to the College of Human Medicine through the American Medical College Application Service (AMCAS) as a Combined Medical Degree/Ph.D. type applicant.
  • Take (or retake, if necessary) the Medical College Admissions Test (MCAT) by the given deadlines.
  • For optimal consideration, application materials including the AMCAS application, MCAT and letters of evaluation should be completed early.

Applicants must pay the AMCAS application fee and the College of Human Medicine Secondary Application Fee. The College of Human Medicine Secondary Application Fee will be waived if the applicant holds an AMCAS fee waiver.

Letters of evaluation

All applicants to the joint M.D./Ph.D. program are expected to submit a minimum of four letters to MSU College of Human Medicine through the AMCAS letters service. Applicants should request three letters of recommendation for the M.D./Ph.D. program component from individuals who can attest to the applicant’s qualifications for the M.D./Ph.D. program, as well as their motivation and ability to conduct research. To address the M.D. component more specifically, applicants should submit at least one letter that reflects their medical/clinical interest and experience and can address personal characteristics consistent with an excellent physician.

The goal of the joint M.D./Ph.D. program is to interview all applicants by the end of October. Applicants are interviewed separately for each program by members of the respective interview panels. The College of Human Medicine Office of Admissions schedules the M.D. interview. Van Andel Institute Graduate School interviews are scheduled through its admissions office. Every effort will be made to schedule the interviews for each program on back-to-back days.

During the College of Human Medicine interview, the applicant will be interviewed by one faculty member and one College of Human Medicine student, tour the medical school and have an opportunity to meet with appropriate faculty and current students.

During the Van Andel Institute Graduate School interview, the applicant will be interviewed by the Graduate School admissions committee, take a tour of the research facilities and be given the opportunity to meet with appropriate faculty and current M.D./Ph.D. students.

Notification

Applicants will be notified of acceptance decisions as soon as possible. Admission to the College of Human Medicine M.D. program is contingent upon acceptance by the joint M.D./Ph.D. Program Selection Committee.

Van Andel Institute Graduate School and Western Michigan University Homer Stryker M.D. School of Medicine

Western Michigan Homer Stryker M.D. School of Medicine (WMed) and Van Andel Institute Graduate School offer an eight-year program leading to both an M.D. and a Ph.D. degree in molecular and cellular biology.

Dual-degree students will:

  • Complete years one and two of medical school
  • Complete the Ph.D. degree, which takes four years (depending on successful completion of the doctoral dissertation)
  • Return to WMed for years three and four of medical school

Students in the M.D./Ph.D dual-degree program complete all courses and clerkships required for the M.D. degree and also complete the biomedical research education including coursework, laboratory rotations and dissertation research within Van Andel Institute Graduate School.

Students are responsible for  tuition and fees for the M.D. program and are eligible for scholarship assistance. Van Andel Institute Graduate School provides financial assistance through a graduate fellowship during the Ph.D. degree that includes a stipend for living expenses; health, dental, vision and life insurance; a full waiver of tuition for the degree; and $2,500 a year for travel to scientific conferences.

Prerequisite Requirements

Applicants for the M.D./Ph.D. dual-degree program must meet all admission requirements and standards of both the Medical School for the  M.D. degree  and the Graduate School for the Ph.D. degree. Applicants must take the MCAT; the GRE is not necessary.

Application

Applicants for the M.D./Ph.D. dual-degree program should begin with the AMCAS application for medical school admission and should select the M.D./Ph.D. option. They should also contact the medical school’s  Director of Admissions  to discuss their interest and coordinate the additional application steps. Select applicants will be asked to interview with both WMed and Van Andel Institute Graduate School on contiguous days.

Students who successfully complete all of the requirements for the M.D. and Ph.D. degrees participate in graduation ceremonies at the Medical School for the M.D. degree and at the Graduate School for the Ph.D. degree.

Fellowship-Ph.D. Program and Medical Internship Program

The Graduate School offers a fellowship-Ph.D. option for applicants who already have earned their medical degree as well as a specially tailored summer internship program for medical students.

This program links subspecialty training in pediatric hematology and oncology with research training in molecular and cellular biology. The training program for each participant will include all components of Van Andel Institute Graduate School’s doctoral program, including courses and dissertation research, although the timelines may be adjusted to meet clinical responsibilities of the clinical fellowship. Clinical training and research experience may be intertwined during portions of the training period. Details of the programs combining the Graduate School Ph.D. with medical school or the clinical fellowship and residency training are articulated in memoranda of understanding with the partner organizations. These memoranda are available upon request.

For more information or to apply, please call the Graduate School at 616.234.5722 or email  [email protected] .

The Medical Student Summer Research Internship Program offered through Van Andel Institute Graduate School provides opportunities for medical school students (between their first and second year) to engage in biomedical research projects in the Institute’s laboratories. Applications for this paid internship program are typically due in early February for the following summer. For more information, please email  [email protected] .

The VAI-MSU Graduate School Program is a collaborative venture through which graduate students in Michigan State University’s BioMolecular Sciences Program (BMS) can conduct their dissertation research in Van Andel Institute laboratories.

The BMS program is the gateway for first-year graduate students who will subsequently select one of the following degree-granting departments or programs:

MSU Departments:

  • Biochemistry and Molecular Biology
  • Microbiology and Molecular Genetics
  • Pharmacology and Toxicology

MSU Interdepartmental Programs:

  • Cell and Molecular Biology

Join the Program

MSU graduate students interested in rotations in a VAI laboratory should  contact the investigator  directly to arrange a meeting to discuss their interests.

Financial Support

MSU graduate students conducting their thesis work at VAI will be appointed as graduate research assistants through MSU and will be paid the same stipend as their fellow graduate students at MSU. Stipend and tuition will be covered for the full term of the thesis project. Students must meet all the standard requirements of their respective graduate programs for graduation, and they will receive their degrees from the MSU program or department in which they enrolled.

Students can choose to have a VAI investigator as their primary thesis adviser or they can have two co-advisers, one from VAI and one from MSU. VAI investigators serving as primary thesis advisers will have adjunct faculty appointments in one of the participating MSU departments.

Current MSU Students in VAI Labs

Peter Huang

Tim Triche, Ph.D.

Alumni MSU Graduate from VAI Labs

Peter Hsueh

Brian Haab, Ph.D.

Daniel Barnett

Sarah Keaton

Lena Brundin, M.D., Ph.D.

Nanda Kumar Sasi

Michael Weinreich, Ph.D.

Sander Frank

Cindy Miranti, Ph.D.

Arkadeep Sinha

Jelani Zarif

Megan Goodall

Jeff MacKeigan, Ph.D.

Aaron DeWard

Art Alberts, Ph.D.

Chih-Shia Lee

Nick Duesbery, Ph.D.

Charles Miller

Sebla Kutluay

Steve Triezenberg, Ph.D.

Laura (Gier) Lamb

Yue Tingting

Susan Spotts

Katie (Sian) Martin

Ying Chou Chen

fully funded phd molecular biology

Copenhagen Bioscience PhD Programme

Fully-funded four year phd programme in an international scientific environment.

fully funded phd molecular biology

News and Events

Copenhagen bioscience snapshots.

fully funded phd molecular biology

The most recent Copenhagen Bioscience Snapshot took place on Nov 9 at Panum, with a talk by Patrick Bryant . Copenhagen Bioscience Snapshots are series of seminar events organized by Copenhagen Bioscience PhD students since 2019. Find more information about past and upcoming Snapshots on the Seminars page .

CPH Bioscience PhD defenses

Phd gruppebillede

Forty-six Copenhagen Bioscience PhD students have defended their theses so far – including all of the students that started in 2016 and 2017. So far in 2024: Kata Krizic, Henry Webel, Philip Gorter de Vries, Zehra Abay-Nørgaard, Matthias Mattanovich, Paul Cachera and Isidora Banjac. We look forward to celebrating more defenses in the coming months.

Student Publications

First-author publications from CPH Bioscience PhD students since the end of 2023 include: Neq2X7: a multi-purpose and open-source fusion DNA polymerase for advanced DNA engineering and diagnostics PCR (Anja Ehrmann); Global, site-resolved analysis of ubiquitylation occupancy and turnover rate reveals system properties (Gabriela Prus); Concerted SUMO-targeted ubiquitin ligase activities of TOPORS and RNF4 are essential for stress management and cell proliferation (Julio Liu); Combining enzyme and metabolic engineering for microbial supply of therapeutic phytochemicals (Maxence Holtz); Seasonally adjusted laboratory reference intervals to improve the performance of machine learning models for classification of cardiovascular diseases (Victorine Muse); Protocol for EHR laboratory data preprocessing and seasonal adjustment using R and RStudio (Victorine Muse); Genome-wide host-pathway interactions affecting cis-cis-muconic acid production in yeast (Paul Cachera); Mass spectrometry-based proteomics data from thousands of HeLa control samples (Henry Webel); Markers of inflammation predict survival in newly diagnosed cirrhosis: a prospective registry study (Henry Webel); Saccharomyces boulardii promoters for control of gene expression in vivo (Carmen Sands); Molecular dynamics-based identification of binding pathways and two distinct high-affinity site for succinate in succinate receptor 1 / GPR91 (Aslihan Shenol); Probing efficient microbial CO2 utilisation through metabolic and process modelling (Philip Gorter de Vries); Ultra-fast label-free quantification and comprehensive proteome coverage with narrow-window data-independent acquisition (Ulises H. Guzman); Altered glucagon and GLP-1 responses to oral glucose in children and adolescents with obesity and insulin resistance (Sara Stinson); Targeting postsynaptic glutamate receptor scaffolding proteins PSD-95 and PICK1 for obesity treatment (Nicole Fadahunsi); Engineering Saccharomyces cerevisiae for fast vitamin-independent aerobic growth (Anja Ehrmann); LiverZap: a chemoptogenetic tool for global and locally restricted hepatocyte ablation to study cellular behaviours in liver regeneration (Elizabeth Ambrosio); The pAblo.pCasso self-curing vector toolset for unconstrained cytidine and adenine base-editing in Gram-negative bacteria (Ekaterina Kozaeva); A quantitative and site-specific atlas of the citrullinome reveals widespread existence of citrullination and insights into PADI4 substrates (Alexandra Rebak); Biosynthesis of natural and halogenated plant monoterpene indole alkaloids in yeast (Samuel Bradley); FAVA: high-quality functional association networks inferred from scRNA-seq and proteomics data (Mikaela Koutrouli); Mapping mammalian 3D genome interactions with Micro-C-XL (Mariia Metelova).

Our students have contributed to at least 220 scientific publications since the programme began in Sept 2016 – including articles in Nature, Science and Cell.

Keep up the good work, everyone!

CPH Bioscience PhD Legacy

fully funded phd molecular biology

The Copenhagen Bioscience PhD programme recruited 109 international students over seven years (2016-2022). Many are enrolled as PhD students at DTU and UCPH currently, but the programme is no longer recruiting new students. CPH Bioscience PhD alumni have gone on to jobs including group leader, post-doc, consulting associate, patent attorney, scientist in industry – and we are very proud of all their achievements.

  • Department of Biology and Biochemistry

Graduate Programs

  • Prospective Students

Ph.D. Programs

fully funded phd molecular biology

The Department of Biology & Biochemistry offers Ph.D. degrees in Biochemistry and in Biology. The Ph.D. program in Biology has two degree tracks: the Cell and Molecular Biology degree track, and the Ecology and Evolution degree track.

Biochemistry

Faculty and graduate student research focuses on biochemical processes at the subcellular and macro-molecular levels and encompasses a variety of fields and methodologies. Areas of study include macromolecular structure and function as elucidated by nuclear magnetic resonance; X-ray crystallographic and spectroscopic techniques; enzyme reaction mechanisms; genomics; computational methods in molecular biology; computational biochemistry/biophysics; computer-aided drug design; signal transduction; neurochemistry; ion channel structure and function; the role of RNA in molecular evolution; the structure and function of virulence factors; and biotechnology.

  • View requirements

Cell and Molecular Biology

The faculty and students in this program share common interests in understanding the molecular mechanisms which drive both fundamental cellular processes and the developmental processes of morphogenesis, cell differentiation and gene regulation. The strength of the program is the diversity of the biological systems under study, which stimulates extensive exchange and collaboration between the various groups. Faculty expertise spans the disciplines of cell and developmental biology, molecular biology, physiology, microbiology, neuroscience, immunology, and genetics.

Ecology and Evolution

This program blends knowledge and methodology from diverse biological disciplines to better understand ecological and evolutionary processes operating at multiple scales—from molecules to individuals to societies to communities. Current research programs include experimental evolution, evolution of development, evolutionary genetics, behavioral ecology, community ecology and evolutionary bioinformatics in systems ranging from bacteria to ants, from protists to grasses. Faculty conduct studies in natural habitats including the Colorado plateau, and coastal salt marshes, and in artificial systems such as petri dishes and theoretical models.

Financial Assistance

The Department of Biology & Biochemistry believes that high-quality graduate studies require a commitment to high-quality research. As a result, our graduate students receive financial support sufficient to provide a modest standard of living that enables them to make a full-time commitment to their graduate training. Some of the types of financial support available to students are listed below. Additional financial assistance may also be available from the College of Natural Sciences and Mathematics and the University of Houston Graduate School .

Teaching Assistantships (TAs)

TAs are the main source of support for first-year students but are available in subsequent years for students not supported by grant funds. TAs will be provided a salary of $2,194.66/month (U.S. $26,335.92 per year). This level of support is sufficient for international students to obtain an F1 visa.

Research Assistantships (RAs)

RAs are the main source of support for students after their first year in the program. RA support is provided through grants to the lab the student joins to conduct graduate research. RA support is currently $2,194.66/month (U.S. $26,335.92 per year).

Graduate Tuition Fellowship (GTF)

Students supported as TAs or RAs are eligible for tuition fellowships to cover their mandatory tuition and fees. These fellowships provide the students with approximately $20,000/year to pay for mandatory tuition and fees. See more information on eligibility criteria .

Non-resident Tuition Waiver

Out-of-state students and international students employed as TAs or RAs receive a waiver of the additional tuition charged to non-residents.

Presidential Fellowship

All applications submitted for admission to the Biology & Biochemistry Graduate Program are reviewed by our Graduate Recruiting and Admissions Committee. This committee is comprised of a group of faculty from each division of the department. Once admitted to the program, accepted applicants are further evaluated for the Presidential Fellowship. The criteria for evaluation are as follows:

  • Appropriate academic coursework in the major
  • Outstanding grades in previous university coursework
  • Demonstrated research ability or potential for research excellence.

If awarded, the student receives $2,000/per year for the first two years. This fellowship is in addition to the monthly stipend and Graduate Tuition Fellowship given to all students admitted to our graduate program. Students must meet minimum full-time enrollment (9 hours) and a cumulative 3.00 GPA to maintain the fellowship each semester it is held.

Houston Livestock Show and Rodeo Scholarship

This scholarship is awarded to outstanding students enrolled in our graduate program. The maximum award is $2,000/year. Recipients must be Texas residents and citizens or permanent residents of the United States. Students apply annually on the UH Foundation website.

Medical Insurance

In addition to their stipend, graduate students employed as TAs or RAs receive $150/month for health insurance. For more information about the student health insurance plan endorsed by and designed especially for the University of Houston, please see Student Health Insurance .

Cost of Living

Houston has a relatively low cost of living compared to most major U.S. cities and many low-cost apartments and houses are available.

Admission Requirements

The minimum entrance criteria for doctoral graduate studies in the Department of Biology & Biochemistry are as follows:

  • Completion of a baccalaureate degree (B.S.) with a major in Biology, Biochemistry, or an equivalent discipline. You can apply to our programs before you complete your degree, provided you graduate before you enter the program. ( NOTE: A prior M.S. is not a requirement to apply to our Ph.D. programs.)
  • Grade Point Average ≥ 3.0 / 4.0 (overall or for the last 60 hours of coursework completed).
  • The GRE is no longer required by our Ph.D. programs . If you decide to submit GRE scores, UH's Institutional Code is 6870.
  • Informative, coherent and well-written statement of purpose .
  • Three strong letters of recommendation .
  • English Language Proficiency Requirement. All applicants, regardless of citizenship status, must demonstrate proficiency in English to obtain admission. To fulfill this requirement, applicants must satisfy one of the following criteria: a) Bachelor's degree (or higher) earned from a regionally accredited U.S. institution or at an institution at which English is the medium of instruction in the following countries: Australia, New Zealand, South Africa, the Bahamas, the United Kingdom, Ireland, Jamaica, Liberia, Trinidad, the Virgin Islands, Antigua and Barbuda, Dominica, Saint Lucia, Saint Vincent and the Grenadines, Barbados, Grenada, Turks and Caicos, and English-speaking Canadian provinces. b) TOEFL. The minimum TOEFL score required is 79 for the internet-based test. The minimum TOEFL score for the new revised paper-based exam is: Reading 20, Listening 20, and Writing 20. TOEFL scores must be received directly from Educational Testing Service (ETS). UH's Institutional Code is 6870. c) IELTS. The minimum IELTS score required is an overall score of 6.5. The testing agency should mail the official results directly to UH. No electronic IELTS are accepted.  d) Duolingo. A minimum score of 105 is required.

Use the online application to submit all your documents electronically. Your references will be automatically contacted to submit their letters of recommendation. Please follow the instructions on the UH Graduate School Application page.

Tips for Applying

What we look for.

We seek to admit students who show a strong record of academic achievement and a high level of motivation and interest. Your record of academic achievement and ability is conveyed by your transcripts, GPA, and Graduate Record Exam (GRE) scores, as well as your letters of recommendation. Your level of motivation and interest is conveyed by your personal statement and letters of recommendation. We will evaluate your application on the basis of your transcripts, test scores (GRE scores for everyone, TOEFL/IELTS scores for foreign students), your personal statement, and the letters of recommendation.

Transcripts and GPA

Successful applicants to our program generally have GPA's of 3.00 or higher. However, a student with a high GPA and a transcript with lots of non-rigorous courses may not be viewed as favorably as a student with a somewhat lower GPA who has taken courses that are essential preparation for graduate work (such as Genetics, Cell Biology, Biochemistry, Evolutionary Biology, etc.). It is not essential to have all of these" foundation" courses before you start graduate school, but if you do not have most of them, you will not be well prepared for graduate school.

The GRE provides information regarding your overall academic ability. You are more likely to do well on the GRE if you prepare for the exam. Preparation guides and practice tests are available at most college bookstores.

Your school may provide assistance in preparing for the GRE; check with your career or academic counseling office.

TOEFL/IELTS

This exam is required for all applicants who have not obtained a prior university degree from an institution where English is the medium of instruction (see list of exempt countries and English Language Proficiency Requirements ). These scores must be officially reported to the University before we can admit you to the program or offer financial support.

Statement of Purpose

Your personal statement is your opportunity to tell us why you want to join our graduate program and what your long-term goals and interests are. You do not have to know exactly what you want to do, or what scientific questions you want to pursue, but you should tell us what excites your interest or curiosity. Be specific. Your statement is also a chance to discuss any aspect of your application (such as academic history) that you feel warrants further explanation.

If there is a reason for your low GPA (a bad semester due to personal difficulties, for instance), you can provide a brief explanation in your personal statement. High GRE scores can make up for a low GPA, and a high GPA can balance out low GRE scores. In some cases, research experience and strong letters of recommendation can make up for low grades and low GRE scores.

Letters of Recommendation

You will need 3–4 letters of recommendation. Most or all should be from your professors, and the letters should come from people who know you personally as well as your academic performance. Make sure your letter writers know your academic record, reasons for wanting to go to graduate school, and long-term goals.

Contacting Faculty

You can contact individual faculty members in our department whose research is of interest to you, either before or after sending your application. Faculty interests and contact information are available on this Faculty Profiles webpage.

Find more information about the application process on the How to Apply page on the NSM website.

  • Open for Applications: June 1
  • Early Deadline: September 1  (apply by this deadline to guarantee full consideration)
  • Late Deadline: November 1
  • Open for Applications: October 1
  • Early Deadline: January 1  (apply by this deadline to guarantee full consideration)
  • Late Deadline: April 1

For More Information

Contact: Rosezelia Jackson 713-743-2633 [email protected]

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ANU International applicants for PhDs in Biomedical Science and Biochemistry

fully funded phd molecular biology

Are you a high-achieving student interested in pursuing a PhD in molecular and biomedical sciences? Do you want to work with top researchers, in a world-class research facility, in a beautiful city at the bottom of the world? Then check out the Division of Biomedical Science and Biochemistry at the Australian National University.

Our Science : We provide an outstanding research environment with a world-class reputation. We are an interdisciplinary division, with highly integrated labs that work at the forefront of membrane transport processes, host-pathogen biology, drug and vaccine development, immune processes, cancer biology and evolutionary genomics. We research a wide range of topics – from the physiology of intracellular parasites to the assembly of bacterial nanomachines; from the structure and biophysics of membrane transporters to the early evolution of animals. If you are interested in doing a PhD with us check out our Research Groups (listed right), and contact prospective supervisors.

Our Values : Our division includes award-winning supervisors and teachers, and we pride ourselves on providing high-quality supervision to our students. Our PhD students are well supported through internal funding, including for conference travel, and our labs are located in a well-equipped, modern research facility. We have a thriving community of PhD students, and our graduates go on to productive careers in many areas of science and beyond. We value diversity and inclusivity in science and have active policies to prevent discrimination. Our faculty and their research teams are comprised of people from all over the globe and from all walks of life.

Location : The ANU is situated in Canberra, Australia’s capital city. Canberra is a well-resourced regional city, set amongst beautiful mountains and surrounded by eucalypt forests. Bike riding and hiking are everyday activities, and we are a two-hour drive away from both winter snowfields and some of the most beautiful beaches on Earth. Canberra’s birdlife is stunning, and we regularly have to brake for kangaroos on our daily commute. Despite its regional setting, Canberra is a vibrant, multicultural city. There are frequent cultural evenings, festivals, art exhibits, music events, and world-class restaurant and coffee scenes.

The Australian National University: We are a research-intensive university and have an international reputation for research excellence. The ANU is frequently ranked as Australia’s top university, and one of the top 50 universities in the world.

Candidates : Fully funded International PhD scholarships are highly competitive at the ANU. To be a viable candidate you need: (1) excellent undergraduate marks; (2) a Masters by Research with a high mark; or an equivalent Honours thesis; or (rarely) equivalent proof of research experience. (3) While not essential, having published research papers in the past is an advantage.

What do I do? First, contact potential supervisors from the list of staff on the Research Groups above (include your CV and grades, and some details about what it is that attracts you to their field of research). If they are interested in supervising you, you can submit an application to commence a PhD via the online link below. Your application should include a detailed CV, full academic transcripts, the name of your potential supervisor, and a brief description of a research project that you and your potential supervisor have discussed. Entry to the PhD program is open to applicants with a Bachelor degree that have also completed (or are in the process of completing) an Honours or Masters research (as opposed to coursework) degree that comprises at least a half year, full time research component and a thesis (5,000-10,000 words). Applicants with significant research experience and publications may be deemed eligible if their achievements can be justified as equivalent to completing an Honours/Masters degree. As part of your application for undertaking a PhD, you will automatically be considered for a stipend scholarship if you tick the scholarship box in the application. You should know that stipend scholarships for international students are very competitive. Your potential supervisor or the Biomedical Science and Biochemistry graduate convenor, Dr Giel van Dooren, can advise you on how competitive your application is likely to be for obtaining a stipend scholarship. If you have questions about any aspect of the PhD application process, you can contact Giel van Dooren ( [email protected] )

The application form is here , along with general information on how to apply and the details about the Doctor of Philosophy program . But read on now as you need to plan ahead in case you are shortlisted. In brief, your PhD application, due by 15 April (for the mid year round) or 31 August,  requires:

  • Academic transcripts and graduation/completion certificates from your Bachelor and Honours/MSc degree (in its original language and, if needed, a translation), with grading scale information.
  • A current CV, including scholarships or prizes, publications and conference presentations.
  • A research proposal (this is quite short and your potential supervisor can help you with it).
  • Proficiency in English: For students from many countries, a current IELTS or TOEFL certificate is required. Note: You can state this is pending. If awarded a scholarship you must then pass the test.
  • Three referee reports. You need to enter your referee details on the application form. The system then automatically sends them a link to complete an online form. NB: You need to ensure your referees are willing and able to write recommendation letters and do so promptly. Plan ahead and alert them.

Student intake

Project status.

  • Adamska Group - Genomic and evolutionary basis of animal development
  • Altin Group - Tumour immunology and liposome targeting group
  • Behm Group - Nematode molecular biology
  • Brock Group - Membrane Structural & Synthetic Biology
  • Broer Group - Membrane transport and nutrition
  • Callaghan Group - Human disease and membrane transport
  • Chung Group - Biophysics of ion channels
  • Corry Group - Transport proteins and computational biophysics
  • Fahrer Group - Immunology
  • Howitt Group - Transporter structure and function
  • Kirk Group - Membrane transport in parasites
  • Lehane Group - Antimalarial drug action and resistance
  • Leyton Group - Assembly, function, and applications of nanoscale bacterial surface structures
  • Maier Group - Molecular mechanisms of malaria pathogenesis
  • Maleszka Group - From molecules to behaviour
  • Saliba Group - Physiology and biochemistry of the malaria parasite
  • van Dooren Group - Cell biology and metabolism of apicomplexan parasites
  • Verma Group - Bacterial and bacteriophage genetics, and vaccine development

Research centre

  • Division of Biomedical Science and Biochemistry

Project team

Support officer.

  • Giel van Dooren

Acknowledgement of Country

The Australian National University acknowledges, celebrates and pays our respects to the Ngunnawal and Ngambri people of the Canberra region and to all First Nations Australians on whose traditional lands we meet and work, and whose cultures are among the oldest continuing cultures in human history.

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PhD in Biological Science (MRC Laboratory of Molecular Biology)

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The Medical Research Council (MRC) Laboratory of Molecular Biology (LMB) is one of the world's leading research institutes. Discoveries and inventions developed at the LMB such as DNA sequencing or methods to determine the structure of proteins, have revolutionised all areas of biology. Our scientists work to advance the current knowledge of biological processes at the molecular level. This information will help us to understand the workings of complex systems, such as the immune system and the brain, and solve key problems in human health.

Every year the LMB International PhD Programme welcomes 20–30 postgraduate students from universities all over the world to do cutting-edge research. At the LMB, we aim to train the scientific leaders of the future: we seek engaged and motivated students, give them rewarding research projects and provide a supportive environment with access to world-class facilities and experts.

Our building is at the heart of the Cambridge Biomedical Campus, a hub of one of the largest and most internationally competitive concentrations of healthcare-related talent and enterprise in Europe. The LMB building's stunning design is home to a globally competitive research centre with state of the art facilities for making the discoveries of the 21st century.

We have an active student association, founded and run entirely by students, who organise numerous events specifically for postgraduates. Postgraduate students register for their PhD with the University of Cambridge and belong to a Cambridge College.

Admission to the PhD programme at the LMB does not require formal completion of an MPhil or equivalent.

The Postgraduate Virtual Open Day usually takes place at the end of October. It’s a great opportunity to ask questions to admissions staff and academics, explore the Colleges virtually, and to find out more about courses, the application process and funding opportunities. Visit the  Postgraduate Open Day  page for more details.

See further the  Postgraduate Admissions Events  pages for other events relating to Postgraduate study, including study fairs, visits and international events.

Key Information

3-4 years full-time, 4-7 years part-time, study mode : research, doctor of philosophy, mrc laboratory of molecular biology, course - related enquiries, application - related enquiries, course on department website, dates and deadlines:, lent 2024 (closed).

Some courses can close early. See the Deadlines page for guidance on when to apply.

Easter 2024 (Closed)

Michaelmas 2024 (closed), easter 2025, funding deadlines.

These deadlines apply to applications for courses starting in Michaelmas 2024, Lent 2025 and Easter 2025.

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  • Infection and Immunity PhD
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  • Biological Sciences (Infection Biology and Molecular Immunology) by advanced study MPhil
  • Clinical Medicine Wellcome Trust PhD

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Doctor of Philosophy (PhD)

Offered By: Department of Molecular Microbiology and Immunology

Onsite | Full-Time | 5 years

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About the PhD in Molecular Microbiology and Immunology Program

Through a departmental core curriculum as well as research area-specific courses, laboratory rotations, qualifying examinations, mentored research, and professional development, all MMI PhD students are prepared to engage in cutting edge research and scholarship that advances knowledge. MMI PhD students hone their scholarship, research and professional skills. Course and laboratory work can address problems in microbial pathogenesis, immunology, disease transmission, and diseases related to malaria, mosquito and arboviral biology.

Students can choose to complete the traditional MMI PhD program or the MMI PhD program concentration in Rigorous, Reproducible, and Responsible Research Investigation in Immunology & Microbiology (R 3 IM). The concentration in Rigorous, Reproducible, and Responsible Research Investigation in Immunology & Microbiology (R 3 IM) conveys a broad background in immunology and infectious diseases research, with a special emphasis on critical thinking, logic, ethics, and written and oral skills to help graduates become communicators of complex scientific concepts and agents of change in their workplaces and communities.

What Can You Do With a Graduate Degree In Molecular Microbiology And Immunology?

Sample careers.

  • Postdoctoral Fellow, Research Associate
  • Science Communicator or Writer
  • Scientist (academia, industry)
  • Specialist in Science Regulatory Affairs
  • Faculty, Professor
  • Science Advocate (nonprofit agencies)
  • Public Health Service (NIH, CDC, FDA)
  • Science Policy Fellow

Curriculum for the PhD in Molecular Microbiology and Immunology

Browse an overview of the requirements for this PhD program in the JHU  Academic Catalogue  and explore all course offerings in the Bloomberg School  Course Directory .

Current students can view the MMI student handbook on the MMI portal site.

Topic Areas

MMI faculty are recognized experts in a wide variety of infectious diseases research areas, allowing our PhD students to study the biology of disease based on their research interests and career goals. Our PhD students gain a comprehensive understanding of infectious diseases that provides a foundation to launch careers that directly tackle critical public health challenges.  

Topic Areas include: 

  • Bacterial pathogenesis
  • Cell biology 
  • Fungal pathogenesis 
  • Medical entomology/Disease ecology
  • Parasite pathogenesis 
  • Rigor, Reproducibility, and Responsibility in Scientific Practice
  • Tick-borne diseases
  • Vaccine development 
  • Vector biology
  • Viral pathogenesis

Admissions Requirements

For general admissions requirements, please visit the How to Apply page.

Standardized Test Scores

Standardized test scores (GRE) are  optional  for this program. The admissions committee will make no assumptions if a standardized test score is omitted from an application, but will require evidence of quantitative/analytical ability through other application components such as academic transcripts and/or supplemental questions.  Applications will be reviewed holistically based on all application components.

Vivien Thomas PhD Scholars

The  Vivien Thomas Scholars Initiative (VTSI)  is an endowed fellowship program at Johns Hopkins for PhD students in STEM fields. It provides full tuition, stipend, and benefits while also providing targeted mentoring, networking, community, and professional development opportunities. Students who have attended a historically Black college and university (HBCU) or other minority serving institution (MSI) for undergraduate study are eligible to apply. To be considered for the VTSI, you will need to submit a SOPHAS application, VTSI supplementary materials, and all supporting documents (letters, transcripts, and test scores) by December 1, 2023. VTSI applicants are eligible for an  application fee waiver , but the fee waiver must be requested by November 15, 2023 and prior to submission of the SOPHAS application.

viven-thomas-scholars

All full-time PhD students will receive the following support for all years of the program: stipend, full tuition, individual health insurance, University Health Services clinic fee, vision insurance, and dental insurance.

Need-Based Relocation Grants Students who  are admitted to PhD programs at JHU  starting in Fall 2023 or beyond can apply to receive a $1500 need-based grant to offset the costs of relocating to be able to attend JHU.   These grants provide funding to a portion of incoming students who, without this money, may otherwise not be able to afford to relocate to JHU for their PhD program. This is not a merit-based grant. Applications will be evaluated solely based on financial need.  View more information about the need-based relocation grants for PhD students .

Questions about the program? We're happy to help.

Alex Kim , Senior Academic Program Coordinator  Department of Molecular Microbiology and Immunology Johns Hopkins Bloomberg School of Public Health 615 North Wolfe Street, Room E5014 Baltimore, Maryland 21205 Fax: (410) 955-0105

Graduate Funding

The Georgetown Biology PhD program is fully funded; a stipend and tuition waiver will be provided for all five years of the program. Students in the program receive their stipends through a combination of research assistantship jobs, teaching assistantship jobs, university fellowships, and external fellowships. 

Stipend funding

Assistantships.

Many students are funded by paid assistantships for all five years. This means that students work as Research Assistants or Teaching Assistants (sometimes called “Teaching Fellows” in the biology department), depending on the semester. For the purpose of managing working conditions and hours, the graduate school differentiates between work you are doing for a stipend (your “employment”) and work you are doing for your degree and dissertation (your “academics”). Teaching Assistants typically help run recitation or lab sections for a course taught by a professor. Research Assistants usually work in the lab of their PI, and can be asked to do work that is not directly related to their dissertation. Both of these jobs are unionized, and subject to the contract negotiated between Georgetown and the Georgetown Alliance of Graduate Employees . Under this agreement, your employment duties cannot exceed 15 hours per week; the remainder of your time is reserved for your own coursework and dissertation research. 

External fellowships

Students are encouraged to pursue external fellowship funding. Fellowships are prestigious, and provide a stipend that is not dependent on a work, or “service” obligation. This can provide more time to focus on your dissertation research, and more flexibility in pursuing research topics outside of the interests of your PI. 

We encourage you to read about fellowship opportunities; here are some places to start:

  • Johns Hopkins list of Graduate Student Funding Opportunities
  • Pathways to Science STEM Graduate Programs and Fellowships
  • UC Santa Barbara EEMB Diversity, Inclusion and Wellness list of funding opportunities (EEB specific)
  • Científico Latino Fellowship database

The most common fellowship that graduate students in the department have won is the NSF Graduate Research Fellowship . This fellowship provides funding for three years of your dissertation; if you are awarded this or other fellowships before you begin graduate school it will strengthen your application and may allow the department to accept you even if funding is limited. If you reach out to a potential advisor in advance, they may be interesting in working with you to develop a GRFP proposal on research you could do in their lab.

Internal fellowships

The Graduate School provides a number of Dissertation Fellowship Awards each year.  Typically, each award will include two semesters of Thesis Research scholarship support.

These capstone awards are intended for doctoral candidates who are in the final stages of their research and writing. Nominees must have completed all coursework and language requirements for their program, passed their comprehensive examinations, and have an approved thesis proposal on file with the Graduate School by the nomination deadline. Each Main Campus doctoral program may nominate one doctoral candidate for a Dissertation Fellowship. The Committee for Graduate Students and Studies will meet to determine the eligibility of students and help to prepare the nomination packet to send to the Graduate School. Interested students or faculty mentors should contact CGSS for more information. 

Internal Professional Development and Research Funding

Travel funds .

The Graduate School and the department support the professional development of graduate students by providing Conference Travel Grants to doctoral students. These grants may be used toward travel and other expenses associated with attendance at a professional meeting and may be requested in advance. The amount of the individual grants will depend on the nature of the student’s participation and the location of the conference.  Please check the Graduate School’s pages for additional information and deadlines. The department travel grants operate on a rolling basis; contact CGSS for more information.

Dissertation Research Travel Award

The Graduate School provides a number of Doctoral Dissertation Research Travel awards of up to $5,000 each. These awards will support the travel costs of doctoral students engaged in archival or field research outside the United States. Grants will not exceed $5,000; the actual amount awarded will depend on budgets submitted by nominees.  

Nominees must have completed all coursework and language requirements for their program, passed their comprehensive examinations, and have an approved thesis proposal on file with the Graduate School by the nomination deadline. Each Main Campus doctoral program may nominate one doctoral candidate for a Dissertation Research Travel award . The Committee for Graduate Students and Studies will meet to determine the eligibility of students and help to prepare the nomination packet to send to the Graduate School.  Interested students or faculty mentors should contact CGSS for more information.

Initiative for Maximizing Student Development (IMSD)

The Georgetown University Initiative for Maximizing Student Development (GU IMSD) reflects our institutional commitment to diversity and scientific workforce development. Leveraging an interdisciplinary network of more than 60 faculty across seven graduate programs and departments, the GU IMSD program will develop a diverse cadre of scholars prepared for careers in the biomedical scientific workforce.

For more information, please visit the IMSD website

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Biochemistry PhD Program

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The application for fall 2024 is closed.

NIH T32 Sponsored Programs

  • Training Grant in Cellular, Biochemical & Molecular Sciences
  • Graduate Women in Science (GWIS)

The Biochemistry & Molecular Biology (BMB) Program is designed for students interested in obtaining a Ph.D. in Biochemistry. As such, the program represents a group of faculty that may mentor a student pursuing thesis research toward the Biochemistry Ph.D. Students interested in a Ph.D. in Biochemistry should apply to the Biochemistry & Molecular Biology Program for admission to graduate studies at the University of Rochester Medical Center.

Why Choose Our Program?

Chenguang Gong in Maquat Lab

We offer in-depth coursework and diverse research opportunities that focus on understanding the biochemical mechanisms of life’s critical molecular processes.

World-class research in our laboratories exposes our students to a variety of the latest methods for sophisticated biochemical analysis, including mass spectrometry, crystallography, microcalorimetry, surface plasmon resonance, microarrays, fluorescence activated cell sorting, light scattering, and spectroscopic methods (including fluorescence lifetime and energy transfer measurements), as well as modern methods for cell culture, protein purification, genetic analysis, and reconstitution of biochemical complexes and reactions.

The flexibility of our training program allows students to train in a number of exciting research areas, and often allows students to develop highly effective interdisciplinary collaborations, resulting in cutting edge thesis projects.

What Sets Us Apart

curriclum

Information on core courses, electives, lab rotations to guide you through our program

research

We support and guide trainees in several areas of research. Browse research by current program trainees

people

Mentor Relationships

Meet the faculty mentors and student researchers on our program and see photos of our department family

Discover the Biochemistry PhD Program

All Student Tours

What Our Students Say...

Leon

"I chose the University of Rochester graduate program because as an UR undergrad, I had early exposure to the amazing research opportunities and investigators. I knew, for a fact, that I would receive high quality training and experience, making me a competitive candidate in the future. What I enjoy the most about the program is the collaboration and support the BMB program provide for its students. They really go above and beyond to ensure that our needs are met."

Leon Harvey

Upcoming events.

Distinct Roles of Paralogous U2AF2 and TatSF1 Proteins in HIV and Host Gene Expression - Thesis Defense

Justin Galardi PhD Candidate, Biochemistry & Molecular Biology Degree Program

  Thu, May 02 @ 2:00 PM   MC | K-207 (2-6408) Hybrid Event

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Latest News

September 5, 2023 Biochemistry Ph.D Student Madeline Jensen Receives Perfect Score on NIH Fellowship Application

June 9, 2023 UR Center for RNA Biology Members and Alumni Attend RNA 2023 in Singapore

May 19, 2023 A Search Engine for mRNA: Algorithm Identifies Optimal Sequences to Improve COVID Vaccines

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PhD in Biochemistry and Molecular Biology

Gain the experience necessary for the development of independent investigators in various areas of biochemistry and molecular biology.

igpbs-biochem-large

Why Study Biochemistry and Molecular Biology?

In the Biochemistry and Molecular Biology doctoral program, students will learn how to perform novel and fundamental research in biochemistry and molecular biology in a laboratory under the mentorship of an advisor, which is a major requirement of this PhD doctoral program. The student is expected to be trained, during the PhD training, to think critically, communicate effectively and independently perform research. This training is crucial for the success of students in their future professional workplaces, including academia and industry. Graduates from our doctoral program are highly successful in acquiring post-doctoral employment, with several excelling in jobs in the biotech and pharmaceutical industries. Some go directly into faculty positions at area colleges, but most do post-doctoral work at highly-regarded institutions around the country.

Why Study Biochemistry and Molecular Biology at UNMC?

There are approximately 45 students in the Biochemistry and Molecular Biology doctoral program, with 4-8 students graduating each year with a PhD degree. The program provides training for both research-based and education-oriented careers, with the average time to the degree of being five and a half years. Biochemistry and Molecular Biology's primary goal is training future scientists and educators to apply critical reasoning skills to solve problems in modern experimental biology and the classroom.

Program Snapshot

Prerequisites.

Admission to graduate work in Biochemistry and Molecular Biology may be granted to a student with a BS or BA from a recognized college or university. The student should have a comprehensive background in chemistry, general physics, mathematics and general biology. 

We prefer students with one year of general chemistry, one year of organic chemistry, mathematics through integral calculus, and one year of biological sciences. A course in biochemistry is highly recommended but is not required, and courses in analytical chemistry, physical chemistry, genetics, and computer applications are helpful. An opportunity to correct course deficiencies is offered during the first year of graduate study.

Careers & Outcomes

During the course of the PhD training, the student is expected to develop the ability to think critically, communicate effectively and perform research in an independent manner. This training is crucial for the success of students in their future professional work places, including academia and industry.

Graduates from our program are highly successful in acquiring post-doctoral employment, with several excelling in jobs in the biotech and pharmaceutical industries. Some go directly into faculty positions at area colleges, but most do post-doctoral work at highly-regarded institutions around the country, including Harvard Medical School, National Institutes of Health, University of California, Stanford University, Stowers Institute for Medical Research, and the University of Pittsburgh School of Medicine.

Related Programs

Students interested in this program may wish to explore:

Two researchers look at lab results

  • Mon. Apr 29th, 2024

PhD Fully Funded Position in Bee Conservation and Systematics at University of Neuchâtel in Switzerland

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By Agristok

fully funded phd molecular biology

PhD Fully Funded Position in Bee Conservation and Systematics at University of Neuchâtel in Switzerland; We are seeking a highly motivated PhD student to work in bee conservation and systematics, starting 1.09.2024. As a member of the Conservation Biology Lab, you will develop two main axes of research. First you will investigate the effectiveness of conservation measures for selected, national priority species of bees in Switzerland using a robust evidence-based framework. Secondly, you will utilize cutting-edge molecular methods to unravel cryptic diversity and tackle unresolved systematic questions within the Central European bee fauna; this systematic research is expected to improve the monitoring and conservation of wild bees and will incorporate an evolutionary biology component such as host-plant or host-parasites (or host-cuckoo bees) interactions.

We offer an engaging research environment committed to bridging the gap between academic research and practical conservation action. You will have the opportunity to contribute to the development of teaching activities in invertebrate conservation and systematics at the University of Neuchatel. The 4-year project is fully funded by the University of Neuchâtel and the Federal Office for the Environment. Salary (60% PhD student) and social benefits follow the rules of the University of Neuchâtel.

Selection criteria:

  • A Master’s degree or equivalent in Biology or Biodiversity
  • A passion for insect ecology or systematics
  • Field experience
  • Proficiency in molecular methods and/or statistics
  • Excellent communication skills and fluency in English

Starting date: The anticipated starting date is 1st of September 2024, with some flexibility.

To apply: Submit your application as a single PDF, including a motivation letter, a complete CV with a copy of your diploma, and the names of two references. Send your application file before May 15 2024 with the subject ‘’PhD application_ [your name]’’ to [email protected] . Interviews are schedule for May 30th . For any question contact Christophe Praz ( [email protected] ) or Clara Zemp ( [email protected] ).

See More Opportunities Like this here .

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COMMENTS

  1. PhD in Biochemistry and Molecular Biology

    In the Biochemistry and Molecular Biology PhD program, faculty, and students work together to increase knowledge of the biochemical and molecular bases of normal and abnormal cellular processes. Our program trains students to be successful independent scientists and gives them the knowledge, research training, and leadership skills to continue ...

  2. Molecular Biology (fully funded) PhD Projects, Programmes ...

    Fully Funded PhD Scholarship in Centrosome Biology. University of Galway School of Biological and Chemical Sciences. Applications are invited from suitably qualified candidates for a 4-year, full-time funded PhD scholarship (s) starting in June-October 2024 affiliated to the School of Biological and Chemical Sciences at the University of Galway.

  3. Fully Funded PhD Programs in Biology

    University of Rochester, Ph.D. in Biochemistry and Molecular Biology (Rochester, NY): The Program is fully funded with a tuition fellowship, competitive stipend, and health insurance. University of Southern California (Los Angeles, CA): PhD students receive full funding throughout their studies, including stipends, and health and dental benefits.

  4. Human Genetics and Genomics, PhD

    The overall objective of the Human Genetics program is to provide our students with a strong foundation in basic science by exposure to a rigorous graduate education in genetics, genomics, molecular biology, cell biology, biochemistry and biostatistics as well as a core of medically-related courses selected to provide knowledge of human biology in health and disease.

  5. Johns Hopkins Biochemistry and Molecular Biology PhD Program

    Rich History Research training of PhD students and post-doctoral fellows in the Department of Biochemistry and Molecular Biology has been supported by a training grant from the National Institutes of Health's National Cancer Institute since 1975. The program's flexible curriculum allows opportunity to take courses throughout the Bloomberg School of Public Health, which has been rated the ...

  6. Molecular Biology (fully funded PhD for international ...

    Search Funded PhD Projects, Programmes & Scholarships in Biological Sciences, Molecular Biology, fully funded PhD for international students. Search for PhD funding, scholarships & studentships in the UK, Europe and around the world.

  7. Ph.D. Program

    The Biology Ph.D. program is part of the larger Biosciences community at Stanford, which includes doctorate programs in the basic science departments at Stanford Medical School. There are two tracks within the Biology Ph.D. program: Cell, Molecular and Organismal Biology. Ecology and Evolution. (Previously a part of the Department of Biology ...

  8. PhD in Molecular and Cell Biology

    The Ph.D. program in Molecular and Cell Biology (MOCB) combines the substantial faculty resources and research facilities of several departments of the College of Natural and Mathematical Sciences and the College of Engineering at UMBC to offer an interdisciplinary, state-of-the-art educational experience for those desiring advanced training in molecular and cell biology. The program provides

  9. EMBL International PhD Programme

    The EMBL International PhD Programme (EIPP), originally established in 1983, provides PhD students with an excellent starting platform for a successful career in science by fostering early independence and interdisciplinary research.. EMBL is dedicated to promoting excellence in the molecular life sciences throughout Europe.To achieve this goal, we inspire and train talented young scientists ...

  10. Doctoral & Ph.D. Program Overview

    Van Andel Institute Graduate School's Ph.D. in molecular and cellular biology is a research-intensive, interdisciplinary program that prepares students for successful careers as independent investigators. Students receive in-depth training in the latest techniques across fields and disease areas, including epigenetics, genetics, cancer ...

  11. Copenhagen Bioscience PhD Programme

    Fully-funded four year PhD programme in an international scientific environment. News and Events. ... Molecular dynamics-based identification of binding pathways and two distinct high-affinity site for succinate in succinate receptor 1 / GPR91 (Aslihan Shenol); Probing efficient microbial CO2 utilisation through metabolic and process modelling ...

  12. Ph.D. Programs

    The minimum entrance criteria for doctoral graduate studies in the Department of Biology & Biochemistry are as follows:. Completion of a baccalaureate degree (B.S.) with a major in Biology, Biochemistry, or an equivalent discipline. You can apply to our programs before you complete your degree, provided you graduate before you enter the program. (NOTE: A prior M.S. is not a requirement to ...

  13. ANU International applicants for PhDs in Biomedical Science and

    Candidates: Fully funded International PhD scholarships are highly competitive at the ANU. To be a viable candidate you need: (1) excellent undergraduate marks; (2) a Masters by Research with a high mark; or an equivalent Honours thesis; or (rarely) equivalent proof of research experience.

  14. Admissions

    The Program is fully funded with tuition fellowship, competitive stipend and health insurance. Admission to the program is on a competitive basis with 10-15 new students matriculating each year. The Graduate Record Examination (GRE) is optional. Applicants whose native language is not English must demonstrate English proficiency.

  15. PhD in Biological Science (MRC Laboratory of Molecular Biology)

    Our scientists work to advance the current knowledge of biological processes at the molecular level. This information will help us to understand the workings of complex systems, such as the immune system and the brain, and solve key problems in human health. Every year the LMB International PhD Programme welcomes 20-30 postgraduate students ...

  16. Biological Sciences (fully funded) PhD Projects, Programmes ...

    10+ fully funded PhD positions at the IMPRS for Molecular Organ Biology. PhD Studies in the Heart of Europe. The International Max Planck Research School for Molecular Organ Biology (IMPRS-MOB) offers fully funded positions to talented and enthusiastic students holding a relevant Master's degree. Read more .

  17. PhD in Molecular Microbiology and Immunology

    Department of Molecular Microbiology and Immunology. Johns Hopkins Bloomberg School of Public Health. 615 North Wolfe Street, Room E5014. Baltimore, Maryland 21205. Fax: (410) 955-0105. The 5-year PhD program allows students to focus on the cellular and molecular mechanisms that drive infectious and immune diseases, the opportunity to take ...

  18. Graduate Funding

    The Georgetown Biology PhD program is fully funded; a stipend and tuition waiver will be provided for all five years of the program. Students in the program receive their stipends through a combination of research assistantship jobs, teaching assistantship jobs, university fellowships, and external fellowships. Stipend funding Assistantships Many students are funded by paid assistantships […]

  19. Biochemistry & Molecular Biology

    The Biochemistry & Molecular Biology (BMB) Program is designed for students interested in obtaining a Ph.D. in Biochemistry. As such, the program represents a group of faculty that may mentor a student pursuing thesis research toward the Biochemistry Ph.D. Students interested in a Ph.D. in Biochemistry should apply to the Biochemistry & Molecular Biology Program for admission to graduate ...

  20. Cancer Biology

    Our lab is part of the Women's Malignancy Program at the Johns Hopkins School of Medicine. Our research is focused on cancer metastasis. Of all deaths attributed to cancer, 90% are due to metastasis, and treatments that prevent or cure metastasis remain elusive. Emerging data indicate that low oxygen tension (hypoxia), which occurs in most ...

  21. PhD in Biochemistry and Molecular Biology

    There are approximately 45 students in the Biochemistry and Molecular Biology doctoral program, with 4-8 students graduating each year with a PhD degree. The program provides training for both research-based and education-oriented careers, with the average time to the degree of being five and a half years.

  22. Molecular Biology PhD Projects

    PhD in Molecular Biology funding options . The research council responsible for funding Molecular Biology PhDs in the UK is the Biotechnology and Biological Sciences Research Council (BBSRC). They provide fully-funded studentships including a stipend for living costs, a consumables budget for bench fees and a tuition fee waiver. ...

  23. Skybright Educational Consults

    9 likes, 0 comments - sky.bright21 on January 7, 2022: "Fully Funded Offer for a PhD in Cell & Molecular Biology. The Assistantship comes with a full tuition waiver and a monthly stipend ...". Skybright Educational Consults | Fully Funded Offer for a PhD in Cell & Molecular Biology.

  24. MolBio Scholars Program

    Apply The Application Portal Opens: April 26, 2024 Application Deadline: June 5, 2024 The Princeton Molecular Biology graduate program invites prospective students interested in graduate training in molecular and cellular biology to participate in the MolBio Scholars program. Scholars will participate in a fully funded visit to the beautiful Pri...

  25. PhD Fully Funded Position in Bee Conservation and Systematics

    PhD Fully Funded Position in Bee Conservation and Systematics at University of Neuchâtel in Switzerland; We are seeking a highly motivated PhD student to work in bee conservation and systematics, starting 1.09.2024. As a member of the Conservation Biology Lab, you will develop two main axes of research.